Every level above 1 grants a hero 5 skill points to use on hero upgrades.
|Animal Discipline||3||Level 4||Animals units in this hero's stack are more strictly handled and taken care of, giving Animal and Mounted units +5 and +200.
|Biology Expert||3||The hero trains its army on the properties of Biochemical weapons and how to improve their bodies, giving Biological units in the army +2.
|Call of the Wild||8||Level 12||Active Ability: Full action |
|Ranger Training||3||Level 8||The hero trains their units in the art of forest stealth. All units in the stack gain Template:Blue.
|Renewal||4||Heal a friendly Biological or Cyborg unit for +5.|
|Visual Acuity||3||The hero gains one extra hex of range when in cover.|
|War Cry||4||Level 4||Active Ability: Full action |
|Armorer||5||Level 6||All units in this hero's army gain +1.
|Hardiness Training||3||Level 4||All units in this hero's army gain +2 resistance against status effects.
|Incentivize||3||Active Ability: Single action |
|Juggernaut||2||The hero gains Template:Blue, allowing them to walk through obstacles. Doing so destroys the obstacle.|
|Siege Master||5||All units in this hero's army gain +2 damage resistance in all channels while in cover.
|Stubborn||3||Level 4||The hero gains Template:Blue.|
|Nitro Battery||5||Level 8||Active Ability: Single action |
|Backstab||4||Level 6||Able to exploit any and all opportunities, this hero has the Template:Blue, dealing 25% more damage when flanking enemies.|
|Subterfuge||3||Employing subterfuge, the hero gains Template:Blue.|
|Strategic Operations Support||2||Operation strength is increased by +6 on operations that target sectors with this hero in them.|
|Shield Battery||4||Level 4||A state-of-the-art shield battery increases the entire army's defenses. All units gain +1.
|Keen Sight||6||Hero gains Template:Blue, and has Template:Blue when using Overwatch.|
|Deploy Cerebral Control Collars||6||Level 8||Active Ability: Full action |
|Escape Module||3||Level 4||When this unit's drops below 0, they automatically teleport to a random position up to 3 hexes away and heal for 15. Can happen only once per battle.|
- Main article: Celestians#Skills
|Aura of Guidance||4||Level 4||Every unit in the hero's army has 10% increased accuracy.
|Compelling Presence||5||Level 8||Active Ability: Full action |
|Deny the Wicked||4||Level 6||Every unit in the hero's army has 2 damage reduction in all channels against attacks from units with Soulburn.
|Revealing Soul||4||Hero gains Template:Blue, and all units that attack the hero have a strength 8 chance of being afflicted with Soulburn.|
|Parasitic Spray||3||Level 4||Active Ability: Single Action |
|Rapid Movements||2||The hero learns to move rapidly to confuse enemies. The hero gains Skitter, making them harder to hit if they move during their turn.|
|Regenerative Spore Cloud||6||Level 8||Hero gains a regenerative spore cloud, healing itself and adjacent friendly Biological and Cyborg units for +20% of their maximum at the end of each turn in tactical combat.|
|Spore Feeding||4||Level 6||The hero surrounds their army with spores that feed on their unit's kills to regenerate their Hit Points. Biological and Cyborg units in the hero's army regain +15 when they kill a Biological or Cyborg unit within 3 hexes of them.